Inversion in "Only when the virus introduces its nucleic acid into a cell does disease occur"
Given this sentence,
Disease occurs only when the virus introduces its nucleic acid into a cell.
Is the following inversion grammatical?
→ Only when the virus introduces its nucleic acid into a cell does disease occur.
Especially the "does disease occur" part.
I think that "occurs disease " is right. Can you explain a rule about this kind of sentence?
Solution 1:
Your sentence is right.
A simple way of looking at the rule of inversion is thinking about question and sentence patterns in English.
The question pattern is:
Verb (V) + Subject (S)
Some learners actually misunderstand the meaning of "v." In the same way that these questions are wrong and unacceptable:
Went he to school?
Like you strawberries?
Your alternative is also wrong:
Occurs disease?
I pointed out these things because the pattern for inversion is the same pattern for questions. And this pattern is:
Auxiliary Verb + Subject + Main Verb
So,
Only when... does disease occur. = OK
Solution 2:
There are lots of different cases where inversion is permissible. There's a really good list at http://esl.about.com/od/advancedgrammar/a/inversion.htm. There are sentences with the structure that you've exhibited, using "only", on this list. There's more detail available about this particular case at http://www.bbc.co.uk/worldservice/learningenglish/grammar/learnit/learnitv34.shtml.
The key point with inversion is that there are are only a limited number of verbs that can come before the subject when you use inversion - just modal verbs and auxiliary verbs. So, you can invert "does" (that is, put it before "disease", as per your example). But you can't invert "occurs".