What's the difference between “by night” and “at night”?

Oxford Dictionaries say by has the meaning of

6. indicating the period in which something happens: ‘this animal always hunts by night’

Then do we need to say, ‘Why does the sky glitter by night?’ And can’t we say, ‘Why does the sky glitter at night?’?

If we can use both, what’s the difference between the two?

Solution 1:

The expression by night is typically used to contrast someone's nighttime activities to their daytime activities, especially when the nighttime activities are unusual or unexpected. You will see it very frequently in the construction X by day, Y by night. Some examples:

• ...treated it like a greasy spoon by day and a hotel by night
  
• I mean, if it's -- if it's somebody that is -- is living as a man that's a woman or living as a woman that's a man or someone that's a prostitute by night and a -- and a secretary or a lawyer by day.
  
• He's a DJ by night, a graphic artist by day. He's multifaceted.
  
• ...he dwelled, chief of all worldly American expats, hanging out with bullfighters, jai alai players, and ex-Spanish Loyalist guerrillas, awash in drink and worldly women, fishing by day, partying by night, writing the whole time.

On the other hand, at night is much less constrained as to the interpretation. The preferred thing to say will be that the sky glitters at night, since this is the expected activity for a sky. To say that an animal hunts by night uses the by night expression because we normally think of day time as the time where animals should be most active, based on our own experience.

Another useful note is that by night can almost always be replaced by at night without altering the acceptability of the sentence, but not vice versa.

Solution 2:

I would put the difference down to more of a sense of drama with use of by. "The tiger hunts by night" sounds more dramatic than "The tiger hunts at night."

Consider the title of the following film: They Drive by Night, which is a hyped-up way of presenting a movie about truck drivers who are trying to survive in their tough world. Had the film been called "They Drive at Night" it would have sounded pretty ho-hum.