What makes up your "Tatau"?

From what I can gather, the relics only put points towards the dots and when you complete the story and the relics and purchase every skill, then it completes it. As you progress through the story, it unlocks the wave like thing at the bottom above the larger 2 sharks, spider and heron. Then, later the story unlocks the tree and the vines beside it. and the mask demon in the middle. I beat the game and only thing I have yet to unlock is the 2 wings at the top and all the dots that trail up to it. Which means I need all the relics to get the wings up top.

Edit: The blue you circled is in fact, from the main story. also right above the middle with the 2 faces is a tree, the tree is from another main story quest. All the rest which you labeled in red and yellow are from skill unlocks that you buy with skill points. when you spend points in spider, it goes towards spider art work etc. The relics only give those dots that you circled in green. I believe that when you get all the relics/ collectibles that you get the 2 final "wings" up top, As you can see, each dot represents 1 relic. Everything else is via main story/ spent skill points towards the animal type. basically that means each line of dots that lead up to the top Shark picture on each side equals 60 relics. since there are 120 relics in all and 2 sides. The memory cards may also play part in the tattoo. Maybe part of the line up top above the wings.

edit #2. Relics only unlock the dots and the wings at the VERY top above the heron's. I know this because I -maybe- collected 5 relics before I beat the game and everything was colored in besides the dots and the 2 wings up top. Right now I am sitting at 24 and the dots, which I counted are 24, is right above the head of the seagull looking icon just above the wave that you highlighted in blue. Now, you do of course get XP from the relics and you might've spent points before looking at it again and thought that, but I was too lazy to get the relics and focused mainly on the story. I could probably show you a picture? Since you have more relics than I do you can see the difference.

The dots represent the relics, and the wings up top are for getting all the relics. There is one missable relic in part of the story. Apprently it's in the cave your friends are in, I'm not sure. Either way, I guess if you missed that one, you are unable to unlock the wings up top.