Does the Spike Grenades perk proc when grenades are bounced off surfaces?

I recently picked up this grenade launcher, which features the perk Spike Grenades. It also has release the trigger to detonate the grenade.

The perk text is

Grenades fired from this weapon do increased damage on direct hits.

This means that I could bounce the grenade off a wall or the floor before it detonates by impacting an enemy.

Does the Spike Grenades perk still trigger if the grenade strikes a non-enemy before it detonates?

Solution 1:

I gave this a test in the Tribute Hall against the Orge tribute.

A direct hit from launcher to the enemy struck for 10,969 + 3,688 = 14,657

A grenade bounced off the floor into an enemy struck for 10,969

Bouncing a grenade off a surface removes 3,688 damage.

From this I would assume that the Spike Grenades perk only triggers if the grenades leaves the launcher and impacts the enemy directly.

This is inline with Spike Grenades having increased impact damage. There is a discussion on Reddit on the topic.