Play Station 4 Online Co-Op without PS+?

I couldn't find a definitive answer on a quick Google, but can you play Borderlands Handsome Jack collection as an online co-op without a subscription to PS+?

Solution 1:

Yes you need a PS+ subscription to play online on the PS4.

Source (Discussion about Handsome Jack collection on PS4)

Solution 2:

A PS+ subscription is generally required to play online, however "free to play" type games are exempt from this rule. Currently there are only a handful of games; Warframe, Airmech, PlanetSide2, etc. Please note that PS+ provides more than multiplayer and those features are still disabled for the previously mentioned games.